<h2 class="nobreak" id="CHAPTER_XIX">CHAPTER XIX</h2>
<h3>THE AGE OF REPTILES</h3></div>
<p>We have already said that in the many hundreds
of thousands of years which went by during
Carboniferous times the sea sometimes advanced
and sometimes receded, and nothing shows this
better than the great thickness of the deposits in which
the coal lies in seams. In America, as in Europe, Asia,
Africa, Australia, and New Zealand, South and Central
America, the Carboniferous system is found. In Arkansas,
in North America, the coal measures attain the remarkable
thickness of 18,000 feet; in the Wasatch
Mountains the Carboniferous strata have been estimated
to be 13,000 feet thick, and in silver-bearing Nevada
10,000 feet. The formations of the Western European
coal measures, like those of Eastern North America,
consist principally of shales and clays, with smaller
amounts of sandstone and limestone. They attain great
thickness, and, including 5500 feet of the Millstone Grit,
are 13,500 feet thick in Lancashire and several thousand
feet thick in many parts of Great Britain and Ireland.
The extraordinary thicknesses show that near our islands
must have been a very extensive and lofty area of land.
In Germany the same strata, thickly seamed with coal,
<span class="pagenum" id="Page_227">-227-</span>
are 10,000 feet thick. There must also have been considerable
volcanic or earthquake action as we know,
because in Germany, near the Hartz Mountains and elsewhere,
there are many igneous rocks thrust into the
strata, and also because in Belgium and in France the
coal strata are very much twisted and contorted. The
same or similar beds are found in Siberia, in Japan, and
in China, where the coal beds are said to be thicker
than anywhere else in the world. The Carboniferous
system is also found in Africa, in the north, south-east,
and south of the continent; and in Australia and New
Zealand Carboniferous strata to the thickness of 10,000
feet are indicated.</p>
<p>At the close of this period the changes ever taking
place transformed the conditions of life in a way the
reverse of that which we have hitherto been examining.
So far each age has shown an increase of life on the age
preceding it. But when the great outburst of carboniferous
activity began to wane it was followed by a
lessening of the wave of life. As we said in the last
chapter, there lie on the top of the coal-bearing strata
beds to which the older geologists gave the name of New
Red Sandstone. But in 1841 the New Red Sandstone
was divided into two distinct geological formations. To
the lower and older part Murchison gave in 1841 the
name of <span class="smcap">Permian</span> (from <i>Perm</i> or <i>Permia</i>, an ancient
kingdom in Russia, where red sandy rocks of this age
form nearly all the surface), but in Germany it is more
frequently called the <span class="smcap">Dyas</span> (from Lat. <i>duo</i>, two), because
in that country it is composed of <i>two</i> well-marked
<span class="pagenum" id="Page_228">-228-</span>
divisions—red sandstones below and magnesian limestones
above. Thus there are two types of this Permian
formation: the Permian type proper—a mixed series of
red sandstones, marls, shales, and limestones, with some
thin beds of coal, as found in Russia; and the Dyassic
type, as seen in Germany.</p>
<p>We do not know what brought about the change:
though we do know that during it there was great
volcanic activity over Europe and that the waning
forests of vegetation and the steaming swamps gave
place to desert plains. Vegetation sank lower and lower.
The forests disappeared or dwelt only in clusters. The
soft sappy trees gave place to hardy pines which clung to
the plains and the mountains, and other sterner types
began to appear, allied to the spruces, yews, and ginkgo.
The ginkgo tree is one of the oldest of the tree type which
now has a living representative. Any reader interested
enough in the matter to walk along Royal Hospital Road,
Chelsea, will find a ginkgo tree just outside the Old Physic
Garden; and of course a good many other examples are
preserved in botanic gardens. The cycads, offshoots of
the ferns, through the strange group of trees known as
the cycado-filices, spread through the woods.</p>
<p>The trilobites, to turn to animal life, all but disappeared,
though one elegant example remained; the
corals were changing; and the lamp-shells were dropping
out. There were a few new species of fishes, but none of
any great strength or capacity, and all preserving still the
tail which is part of the backbone. There was an all-round
impoverishment of life—so great, indeed, that the
<span class="pagenum" id="Page_229">-229-</span>
early geologists used to believe that a complete destruction
of all life followed this, the closing stage of the Palæozoic
era, and that a re-creation followed. They would have
been confirmed in this belief had they but known that
intense cold and glaciation were setting in where the
tropics were situated and that the dryness of vast deserts
was sweeping away life elsewhere. But we now know
that life was not entirely lost; that many species survived,
and that others, altering to suit altering conditions,
became stronger in the process. Nevertheless, life was
greatly impoverished. A census made a few years ago
gave the known animal species of the Carboniferous
period as 10,000, while those of the Permian period were
only 300, or three per cent. Possibly the percentage was
larger than that, but still it was small.</p>
<p>But if the Permian was poor in life it was very interesting.
The amphibians had been growing in strength
during the later stages of the Carboniferous age, and may
possibly have been more numerous then than at any
other time, for the vast swamps were very favourable for
them. They diminished in the Permian, though the
Permian amphibians showed some advances, and began to
assume a likeness to reptiles. Perhaps the reptiles may
have first appeared in the Carboniferous, but they declared
themselves in the Permian age. Two great branches
of reptiles seem already to have defined themselves;
perhaps they had never formed a common group as
reptiles, but had separated while still amphibians. The
one bore resemblance to, and were perhaps the forerunners
of, the great hosts of lizards, crocodiles, dinosaurs,
<span class="pagenum" id="Page_230">-230-</span>
ichthyosaurs, and flying saurians, which are the most
pronounced of the reptiles, and of which we shall have a
great deal to say presently. The other group were perhaps
the ancestors of the turtles and plesiosaurs which
appeared later, and possibly led the way to the mammals.
This rapid and diverse spreading out of the reptiles in
a period when life as a whole was at a low ebb is not a
little remarkable. These creatures seem to show the
arrival of a more pronounced form of <i>air-breathing</i>
animal; and that may have been the consequence of the
presence of more oxygen in the earth's atmosphere. Of
the Permian amphibians, one of the most interesting was
like the Sphenodon, which still creeps about the northern
islands of New Zealand. The most striking of the
reptiles was the <i>Naosaurus</i>, a beast-like creature with a
high back of spines webbed together like a solid porcupine.
It was from three to ten feet in length.</p>
<p>All these changes were brought about by the general
withdrawal of the sea, both in the North American continent
and in Europe. In both continents there are beds
which accumulated fresh water; in both beds which were
laid down in salt lakes or inland seas; and in both beds
which were laid down on the floor of seas washing the continents.
Great areas seem to have been sometimes dry
and sometimes submerged; other and greater areas,
bordered by ice and sometimes swept by icy blasts, or
subject to burning sun in summer, were deserts such as we
are aware of now in Asia or North Africa or mid-Australia,
but much larger in extent than any of these.
It was in conditions such as these that the ancient
<span class="pagenum" id="Page_231">-231-</span>
or Palæozoic rocks came to an end and the Mesozoic
or Middle Period began.</p>
<p>The Middle period of strata and of the life which those
strata have preserved has usually been separated from the
older rocks because, owing to the great period of arid
desert conditions, the character of life changed a great
deal; but fuller knowledge shows that the links were
still there, and that ceaseless adaptation of animals to
their surroundings was ceaselessly going on. We need not
follow closely all the changes and relationships, and only
much greater knowledge than geologists yet possess will
enable them to trace all the alterations of the land and sea.
But we may trace the alterations in the appearance of the
continents in broad outline. Nearly the whole extent of
the British Isles was now above the sea, and was enjoying
a climate perhaps as cold as present-day Iceland. To the
south and east of Scotland was a great shallow inland lake,
while north of Great Britain a huge plain stretched across
Europe. To the south of the lake was a belt of land,
and farther south still the sea had invaded Italy and
reached to Southern Germany, and in this sea was being
laid down the limestone which in later eras was to be
elevated into the mountains of the Apennines, the Alps,
and the Pyrenees. North Africa was under water, but
farther south the uplifted lands were joining hands with
India. Sea swept part of Asia, but North America was
larger and broader than it is now, her western coast
stretching farther out into the ocean.</p>
<p>In this period, which is called the Triassic (the name
given to it by the German geologist Bronn because of the
<span class="pagenum" id="Page_232">-232-</span>
three distinct beds he found in it, though the middle of
these, a shelly limestone, does not exist in Great Britain),
there was a wide development of large reptiles and
amphibians. We cannot enumerate them all, for not one
chapter nor one volume would suffice to deal adequately
with the reptiles of the Trias formations and of the
Jurassic rocks which followed them, and of the Permian
which preceded them. But we may speak of some of
them. One of the most striking was the <i>Pareiasaurus</i>,
which has been found in the Jurassic sandstones and limestones
of South Africa, of Russia, of India and Scotland,
and of middle England. The skeleton of the Pareiasaurus
looks like that of a gigantic pug dog eight feet
long; but it was a comparatively harmless animal, the
teeth of which show that it largely fed on vegetable
food.</p>
<p>In Sir E. Ray Lankester's lectures on "Extinct
Animals" he described the finding of a great many
of these fossil reptiles by Professor Amalitzky on the
banks of the River Dwina, near Archangel. There is
a cliff of Permian strata on the banks of the Dwina,
and in this cliff there is a peculiar pocket or accumulation
of sandy matter with large hard nodules embedded
in it. These nodules are removed and broken up for
mending the roads. The pocket seems to be in a fissure
and of Triassic age, later, that is to say, than the
Permian rocks on either side of it. However that may
be, the nodules are usually removed for road-mending,
and four or five years ago Professor Amalitzky on visiting
the spot was astounded and delighted to find that
<span class="pagenum" id="Page_233">-233-</span>
when broken each nodule was seen to contain the skeleton
or skull of a great reptile. The Russian geologist
determined to make a most thorough investigation of
this wonderful deposit, and for years spent several thousand
pounds in having the nodules dug out by the
peasants after the year's farming work was over, and in
removing them to the University of Warsaw, where with
the finest instruments and greatest care the nodules are
opened, and each bone removed in fragments is put
together from its more or less broken parts, and firmly
cemented and set up in its natural position as a complete
skeleton.</p>
<p>These Pareiasaurs reconstructed by Professor Amalitzky
were about as big as well-grown cattle, but not
so high on the legs. Living at the same time, and its
skeleton now found near them, was an enormous and truly
terrible flesh-eating animal, with a skull two feet long
and enormous tiger-like teeth. This creature was named
Inostransevia. No doubt the vegetarian herds of Pareiasaurus,
whose small peg-like teeth indicate their harmlessness,
were preyed on by the terrible Inostransevia, as
were their brethren in South Africa devoured by other
carnivorous reptiles of that remote Triassic age. So we
see the co-existence of blood-sucker and victim—of the
destructive oppressor and the helpless oppressed—forced
on our attention in these two localities, Russia and South
Africa, in days long before man was. Other land forms
were grotesque and curious in shape, the Chelonians for
example, big birds and crocodiles rolled into one, and
clothed in lizard-like skin—queer pear-shaped brutes with
<span class="pagenum" id="Page_234">-234-</span>
huge hind limbs, short fore limbs, narrow chests, and
pigmy skulls.</p>
<p>Both branches of the reptilian horde, those representing
the saurians and those which were the forerunners of the
mammals, sent delegations to the sea before the close
of the Triassic period. The Ichthyosaurs represented
the more pronounced reptilian line; the Plesiosaurs were
the representatives of the coming mammals. It is not
difficult to find good reason for this movement to the
sea. Besides the inevitable tendency of every masterful
race to invade all accessible realms, the renewed extension
of the sea that set in during the Triassic period and
became pronounced before its close, especially invited
this, for the shallow waters creeping out upon the land
with their now prolific life set tempting morsels before
the voracious reptiles, on the one hand, while on the
other, the reduction of the land area and the restriction
of their feeding-grounds, intensified by the multiplication
of the reptiles themselves, forced a resort to the sea.
One of the reptiles of this period, the <i>Lariosaurus</i>, shows
by its development how the change affected the reptiles.
In the earlier stages of the Trias it resembled a rather
swollen alligator with four limbs symmetrically situated
and used for crawling. In the later forms of these
reptiles the limbs were modified with paddles, and all
power to move about on land was lost.</p>
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